RETURN TO DATABASE - ALTERNATIVE CLASSIFICATION
SALTICIDAE OF THE WORLD - Abstract - INDEX of Genera - FOREWORD & KEY to groups of genera! - AEURILLINES - Comparison HISTORICAL classification of Aelurilleae - AMYCINES- AMYCOIDA VARIA - ASTIAINES - BELIPPINES - CHRYSILLINES - COCALODINES - COLONINES [=Thiodinines] - DENDRYPHANTINES - DIOLENINES - EUOPHRYINES-PART 1 Introduction-Donoessus- EUOPHRYINES-PART 2 Echeclus-Pystira - EUOPHRYINES- PART 3 Rhyphelia-Zenodorus - EUPOAINES - EVARCHINES - FOSSILS - HABRONATTINES - HARMOCHIRINES - HELIOPHANINES - HISPONINES - HYLLINES - ICIINES - LAPSIINES - LIGONIPEINES - LYSSOMANINES - MENEMERINES - MYRMARACHNINES - NOTICIINES - PELLENINES - PSEUDICIINES - SIMAETHINES - SITTICINES - SPARTAEINES - THIRATOSCIRTINAE - YAGINUMAELLINES - YLLENINES - UNCLASSIFIED TEMPORARY - Maddison's views on Salticidae phylogeny 2014 - Omoedus synonymy
OUT OF DATE CLASSIFICATORY SYSTEMS -Simon's classification - Petrunkevich synthesis-1928 - Bonnet's list of subfamilies - Prˇszynski's revision of subfamilies 1976.

Monograph of the Salticidae (Araneae) of the World 1995-2015. Introduction to alternative classification of Salticidae
Classification of the Salticidae genera of the World
group of genera
HELIOPHANINES
by Jerzy Prószyński

Version December 31st, 2015
Note on illustrations. This introduction to classification of Salticidae includes only species defined by diagnostic illustrations in the taxonomic literature, arranged by similarities of embolus and spermophor. Images are displayed as thumbnails to save space and allow for rapid scanning of theoretically closely-related species. Thumbnails are small versions of illustrations in other parts of the monograph, whera are displayed together with particulars of their sources and permissions.

Gen. Heliophanus Koch C.L., 1833 (143 species)

Type species Heliophanus cupreus.
Drawings below are integral part of definition.

Subgen. Heliophanus (Heliophanus) Wesołowska, 1986 (81 species)

+
Heliophanus (Heliophanus) cupreus: Zabka M. 1997. Fauna Polski 19: 5-187, figs 148-156 (Type Species). ©Photo A. Senglet.
b)+
Heliophanus (H.) aeneus +b) apiatus : Zabka 1997. F. Polski 19: 5-187, f 132-139.+ ©photo B. Knoflach
+b) Wesolowska 1986. Ann. zool. 40, 1: 10, 39-40, ff. 447-455, + ©Photo J. Lissner. By courtesy.
Heliophanus (Heliophanus) curvidens : Proszynski 2003a. Annales zoologici: 72-73, figs 269-272
+b) ©Photo Antoine Senglet. By courtesy.
+
Heliophanus (Heliophanus) auratus : Wesolowska W. 1986. Annales zoologici, 40, 1: 10, 212, f 629-639 + Zabka M. 1997. Fauna Polski 19: 56, f 140-147 + ©Photo Antoine Senglet. By courtesy.
>
Heliophanus (Heliophanus) melinus : : Wesolowska W. 1986. Annales zoologici, 40, 1: 220, f 741-751. + © Photo A.Senglet + J. Lissner. By courtesy.
Heliophanus (Heliophanus) mordax : Proszynski. Ann. zool. 2003a: 53, 1: 79, f 303-308 + © Photo A.Senglet. By courtesy..
b)+
Heliophanus (H.) activus +b) agricola : Wesolowska 1986. Ann. zool., 40, 1: 11, 224, f. 795-802 + 10, 209-210, ff. 589-600
+ Wanless 1984c. Ann. Mus. Roy. Afr. Cent., 241: 70-73, f 23a-g + ©Photo Antoine Senglet. By courtesy.
+
Heliophanus (H.) agricoloides : Wunderlich J. 1987: 277-278, f 695-698. + ©Photo J. Lissner.t. By courtesy.
b)
Heliophanus (H.) auratus + aviculus :Zabka 1997. F. Polski 56, f 140-147 + ©Photo Antoine Senglet +Wesolowska 1986.
Ann. zool., 40, 1: 9, 29, ff.` 279-288. By courtesy.
>
b)c)
Heliophanus (H.) baikalensis +b) butemboensis +c) conspicuus : Proszynski. 1979. Ann. zool., 34: 308, ff. 98-100
+ Wesolowska 1986. Ann. zool., 40, 1: 1: 10, 45, ff. 536-537 + Marusik, Cutler 1989. A. arachn. 37: 54, figs 4-7
+b) Wesolowska 1986. Ann. zool., 40, 1: 8, 24, ff. 199-200 +c) Wesolowska 1986. Ann. zool., 40, 1: 10, 41-42, ff. 487-491,
901-map.10. By courtesy.
Heliophanus (H.) creticus: Wesolowska 1986. Ann. zool., 40, 1: 10, 40-41, f 467-475. By courtesy.
>+
b)+ c)
Heliophanus (H.) dampfi + camtschadalicus + deamatus : Proszynski J. 1979. Ann. zool. 34: 309, f. 103 + 309, f. 103
+ Zabka 1997. F. Polski 19: 5-187, figs 157-165 + ©Photo Photo J. Lissner +b) Wesolowska, Marusik 1990: 91, f. 1-5
+ Peckhams 1903. 14 (1): tab. 20, figs 3a-c. By courtesy.
Heliophanus (H.) decoratus: Proszynski 2003a. Annales zoologici. 53 (1): 73-75, figs 273-280. By courtesy
+
Heliophanus (H.) dubius : Zabka M. 1997. F. Polski 19: 5-187, figs 166-173.+ ©Photo J. Lissner. By courtesy.
b)
c)
Heliophanus (H.) dunini +b) koktas +c) falcatus: Rakov, Logunov 1996a. Arthr. Sel. 5 (3/4): 80-82, figs 51-54,
59-60, 61 +b) 89, figs 87-96. +c) Wesolowska 1986. Ann.s zool., 40, 1: 9, 35, f. 367-375. By courtesy
+
Heliophanus (H.) dux :Wesolowska, van Harten 1994. Yemeni-Germ. Plant Prot. Proj. 86: 35, fig. 75-83. By courtesy
Heliophanus (H.) encifer: Proszynski 2003a. Annales zoologici. 53 (1): 77, figs 287-291. By courtesy
+
Heliophanus (H.) equester : Proszynski 2003a. Annales zoologici.: 77-78, f 292-295 + ©Photo A. Weinstein. By courtesy.
+
Heliophanus (H.) flavipes : Zabka 1997. F. Polski 19: 5-187, figs 174-181 + ©Photo A. Senglet + J. Lissner. By courtesy.
+
Heliophanus (H.) cassinicola : Wesolowska 1986. Ann. zool., 40, 1: 8, 28, ff. 261-278, 917 + ©Photo B. Knoflach. By courtesy.
b)
Heliophanus (H.) forcipifer +b) gladiator :Rakov, Logunov 1996a. Arthr. Sel. 5 (3/4): 88-89, figs 61, 76-83
+b) Wesolowska 1986. Annales zoologici, 40, 1: 10, 38, ff. 434-437. By courtesy.
b)c)d)
Heliophanus (H.) glaucus +b) hamifer +c) harpago +d) ibericus : Wesolowska 1986. Ann. zool., 40, 1: 10, 208, ff. 558,
567-569 +b) 9, 37, ff. 420-431 +c) 9,33, ff. 341-345 +d) 10, 209, ff. 585-588 By courtesy.
b)c)
Heliophanus (H.) imerinensis +b) improcerus +c) innominatus : Wesolowska 1986. Ann. zool., 40, 1: 9, 37, f 412-419
+b) 9, 35, ff. 376-383, 921 +c) 9, 38, ff. 432-433. By courtesy.
b)
c)d)
Heliophanus (H.) kankanensis +b) lawrencei +c) mauricianus +d) maralal[?] : Wesolowska 1986. Ann. zool., 40, 1:
9, 36, ff. 384-400, + 9, 34, ff. 359-366, +9, 36, ff. 401-411 +d) Wesolowska 2003. 14(2): 272-273, figs 73-74. By courtesy.
Heliophanus (H.) kochii :Wesolowska 1986. Ann. zool., 40, 1: 11, 217, f 702-717 + ©Photo Senglet + J. Lissner. By courtesy.
+
Heliophanus (H.) lineiventris : Proszynski 1982.. Ann. hist. nat. Mus. hung., 74: 1982: 283-284, ff. 28, 31, 35
+ ©Photo J. Lissner. By courtesy.

b)
Heliophanus (H.) montanus +b) ndumoensis :Wesolowska 2006b. Afr. Ent., 14 (2): 233-234, f 38-39
+b) Wesołowska, Haddad 2013: 54 (1): 205-206, f 12, 82–85. By courtesy.
+
Heliophanus (H.) orchesta : Wesolowska 1986. Ann. zool., 40, 1: 9, 30, ff. 295-309 + ©Photo T. Szuts. By courtesy.
+>
Heliophanus (H.) parvus: Wesolowska, van Harten 2002. F. Arabia 19: 379, f 33-37 + ©Photo B. Knoflach. By courtesy
+
Heliophanus (H.) patagiatus Wesolowska 1986. Ann. zool., 40, 1: 11, 221, ff. 752-759 + ©Photo B. Knoflach. By courtesy.
+b)
Heliophanus (H.) potanini +b) pratti : Proszynski 1982. Ann. hist. nat. Mus. hung. 74: 284-285, ff. 29, 32
+ Wesolowska 1981. Ann. zool., 36, 1: 133-134, f 14-17 +b) 1986, 40, 1: 10, 39, ff. 442-446
b)c)d)
Heliophanus (H.) ramosus +b) semirasus +c) robustus +d) comorensis :Wesolowska 1986. Ann. zool., 40, 1: 10,
42, ff. 503-506 +b) 9, 34, ff. 354-358 +c) Wesołowska, Russel-Smith 2011. Ann. zool. 61(3): 573-574, f 69-74, 225
+d) Dierkens, 2012: 72-73, f 7-11. By courtesy.
+
Heliophanus (H.) rufithorax :Wesolowska 1986. Ann. zool., 40, 1: 11, 218, ff. 718-725 + ©Photo A. Senglet. By courtesy.
+
Heliophanus (H.) simplex :Wesolowska 1986. Ann. zool., 40, 1: 10, 210, ff. 601-611 + ©Photo J. Lissner. By courtesy.
+b)c)d)
Heliophanus (H.) sororius +b) splendidus +c) tristis +d) wesolowskae : Wesolowska 2003a. Genus, 14(2): 286-287, f 122-126 + Wesolowska, Haddad C.R., 2014. Lesotho. Afr. Invert. 55 (2), 246-247, f 33 +b) 287-289, figs 127-132 +d) 289-289, figs 133-134. Rakov, Logunov 1996. Arthr. Sel. 5 (3/4): 101, f 61, 140-145. By courtesy
++
Heliophanus (H.) tribulosus :Wesolowska 1986. Ann. zool., 40, 1: 10, 42, ff. 492-502+ ©Photo A. Senglet + J. Lissner.
b)c)
Heliophanus (H.) transvaalicus +b) turanicus +c) excentricus :Wesolowska 1986. Ann. zool., 40, 1: 9, 33, ff. 346-353
+b) Rakov, Logunov 1996a. Arthr. Sel. 5 (3/4): 99, figs 21, 30, 55-56, 132-136 +c) Ledoux, 2007. 17(2): 27-28,
f 25 + Dierkens 2012. 71, 4-6. By courtesy.
>>b)
Heliophanus (H.) ussuricus + verus: Proszynski 1982. Ann. hist. nat. Mus. hung. 74: 285, f 30 + 1979 Ann. zool., 34: 310,
+ © Ono, Ikeda, Kono 2009: 572. + Wesolowska 1986. Ann. zool., 40, 1: 11, 224, f. 795-802. By courtesy.
+
b)
Heliophanus (Heliophanus) capicola +b) deformis : Wesolowska 1986. Ann. zool., 40, 1: 11, 225, ff. 807-813 + Wesołowska, Haddad C.R., 2014. Lesotho. Afr. Invert. 55 (2), 237-238, f 24, 25+b) 11, 226, ff. 819-824. . By courtesy
b)c)
Heliophanus (Heliophanus) difficilis +b) africanus +c) feltoni : Wesolowska 1986. Ann. zool.,40, 1: 11, 226, f 825-827
+b) 11, 224, ff. 803-804 +c) Logunov, 2009. Turkey. Contrib. Nat. History 12: 901-903, 1-5, map 1..
b)c)+
Heliophanus (Heliophanus) acutissimus +b) menemeriformis +c) chovdensis : Wesolowska. 1986. Ann. zool., 40, 1:
11, 222, f 762-765 +b) 230, f 866-870 +c)
Proszynski 1982. Ann. hist. nat. Mus. hung.: 283, f. 34 + Rakov, Logunov
1996a: Arthr. Sel. 5 (3/4): 70-70, figs 10-20.
By courtesy.
b)
Heliophanus (Heliophanus) iranus +b) pauper Wesolowska 1986. Ann. zool., 40, 1: 11, 223, f 776-786 +11, 228-229, f 850-851 .
+b)
Heliophanus (Heliophanus) abditus +b) xerxesi : Wesolowska. 1986. Ann. zool., 40, 1: 11, 221, ff. 760-761
+ .Wesolowska, van Harten. UA Emir., 2010: 3: 36-37 +b) Logunov, 2009. Contrib. Nat. History 12: 905-907, 13-17.
+b)
Heliophanus (Heliophanus) gramineus +b) konradthaleri: Wesołowska & Haddad 2013. Afr. Invert.: 54 (1): 204-205,
f 11, 80, 81 +b) Logunov, 2009. Turkey. Contrib. Nat. History 12: 903-905, 6-12. By courtesy.
+b)c)
Heliophanus (Heliophanus) infaustus [s. orchestioides] +b) kittenbergeri +c) vittatus [misplaced]: Wesolowska 2012a.
Afr. zool. 20(2): 328-329 + Wesolowska 1986. Ann. zool., 40, 1: 9, 32, ff. 327-340 + Wesolowska, Cumming 2008. Ann. zool.
58:182, f 44-49 +b) Caporiacco. 1947: fig 68 +c) Denis, 1958. Videns. Meddr dansk naturh. Foren.: 106, f. 35. By courtesy.
+b)
Heliophanus (Heliophanus) lesserti +b) machaerodus : Wesolowska 2011. © Genus, 22(2): 315-316, f 20-24, 88
+ Wesolowska W. 1986. Ann. zool., 40, 1: 9, 31, ff. 320-326 +b) 11, 219, ff. 726-732. By courtesy.
+
Heliophanus (Heliophanus) malus : Wesolowska 1986. Ann. zool., 40, 1: 11, 223, f 787-788,
+ Proszynski. Ann. zool.. 2003a: 79, figs 300-302. By courtesy.
b)+
Heliophanus (Heliophanus) pygmaeus + similior [misplaced] + sp : Wesolowska W. 2004b.15(2): 275-280, f 1-13
+b) Ledoux, 2007. Rev. Arachn. 17(2): 30-31, f 27, 28, 29C +31, f 27c-d, 29e. By courtesy.
b)c)
Heliophanus (Heliophanus) stylifer +b) tenuitas +c) variabilis : Wesolowska W. 1986. Ann. zool., 40, 1: 10, 43, ff. 507-513
+b)Wesolowska 2011. © Genus, 22(2): 318-319, f 28-31 +c) Ledoux, 2007. Rev. Arachn. 17(2): 25, f 25 [sic! =26], 30.
\\\
Heliophanus mucronatus : Wesolowska 1986. Ann. zool., 40, 1: 11, 227, ff. 837-842. By courtesy.