SALTICIDAE OF THE WORLD - KEY to groups of genera! - INDEX of Genera
KEYS TO INDIAN SUBCONTINENT -KEY to groups of genera of INDIAN SUBCONTINENT - INDEX to Indian genera - Aelurillinae of India - Chrysillinae of India - Dendryphantinae of India - Euophryinae of India - Eupoainae of India - Evarchinae of India - Harmochirinae of India - Heliophaninae of India - Hyllinae of India - Iciinae of India - Lyssomaninae of India - Menemerinae of India - Myrmarachninae of India - Noticiinae of India - Pelleninae of India - Pseudiciinae of India - Simaethinae of India - Sitticinae of India - Spartaeinae of India - YAGINUMAELLINAE-India - Ylleninae of India - Unclassified-TEMPORARY of India - REFERENCES to Salticidae of India - Literature quotation proposals.- Maddison's views on Salticidae phylogeny 2014 - Omoedus synonymy
OUT OF DATE CLASSIFICATORY SYSTEMS -Simon's classification - Petrunkevich synthesis-1928 - Bonnet's list of subfamilies - Prˇszynski's revision of subfamilies 1976.
Monograph of the Salticidae (Araneae) of the World 1995-2014
Introduction to classification of Salticidae
a subchapter

A Key to Salticidae of the Indian Subcontinent
group of genera

trial version

by Jerzy Prószyński & John T.D. Caleb
Version June 7th, 2015

Disclaimer. This is a work in progress, continually being developed and revised. Proposed new taxonomic names, included here are not valid unless subsequently published in accepted taxonomic papers. Note on images. This introduction to a classification includes only species defined by diagnostic illustrations in the taxonomic literature and Internet, arranged by similarities of embolus and spermophor. Images are displayed as thumbnails to save space and allow for rapid scanning of supposedly closely-related species. Thumbnails are small versions of illustrations in other parts of the monograph, where are displayed together with particulars of their sources and permissions.


Type genus Yllenus.
(albocinctus group) (arenarius group) (hamifer group) LOGUNYLLUS MARUSYLLUS Yllenus


YLLENINAE are characterized by presence of scopula on tarsi I-II - brush of setae stretching from claws onto ventral surface, an adaptation for quick submerging in sand, where these spiders make nests, retreats and produce cocoons (Bartos 2000, 2005). Caparace highest at eyes III, from where thorax is gently sloping. Male palp robust, with large "conductor" process splitting off the basis of embolus, embolus is hair like. Palpal femur broad, crescent shaped. Epigyne with prominent median pocket, spermathecae and copulatory ducts either heavily sclerotized or, if thinner, then ducts form of complicated coils, in some species making double spiral. Chelicerae without any retromarginal teeth, prolateral edge with sclerotized rim, often protruding as small tooth distally, near fang.
REMARK. YLLENINAE are ground dwellers, living on sands in Temperate Asia, N Africa and Europe (South and Central). Combinations of their characters is unknown in other subfamilies, including ground dwelling AELURILLINAE. The structure of this taxon fit definition of a subfamily (ranked group of genera in this paper), with several mutual characters, including appearance and biology. Male palps, epigyne and internal structure of epigyne delimit three groups of species, here described as three genera.
Historical note. The main concept of the subfamily was developed during basic, extensive research by Prˇszynski 1968e, in which he presented modern definition and contents of the genus Yllenus and proposed its division into three species groups, now proposed further to become full genera. The scope of the genus was considerably enlarged by large number of new species, and details of distributions of the previously known ones, by Logunov and Marusik 2003b. These data were later summarised in the database of Salticidae (see latest version 2015).

Characters of Ylleninae: a) embolus, conductor and bulbus + b) epigyne, spermathecae and ducts, carapace
posterior slope, scopula and cheliceral dentition
: a) SEM by Logunow and Marusik 2003b: 15, f 53-56
+b) Proszynski J. 1968e. Annales zoologici, 26: 409-494, f 1-185.

Guide to identification of genera by palps and internal structure of epigyne

Guide to identification of genera by epigyne

Guide to identification of genera by color pattern

Gen. LOGUNYLLUS - gen. n.
expected but not yet reported from Indian Subcontinent

[name unpublished, proposed as partial synonym for the genus Yllenus (in part - albocinctus group Prˇszynski, 1968)].

Type species Yllenus (albocinctus group) albocinctus Simon, 1876
ETYMOLOGY. Named in honor of prominent arachnologist - Dr. D.V. Logunov [LOGUN-ov-YLL-en-US], author of a large number of publications on Salticidae, including big monograph of the genus Yllenus. Grammar gender - male.
DESCRIPTIONS. Palps, epigyne and internal; structure of epigyne relatively less specialised than in two remaining genera (see drawings below).

Yllenus (albocinctus group) albocinctus : Proszynski J. 1968e. Ann. zool. 26: 463-466, f 25, 38, 52, 66, 136-143
+ Proszynski 1982. from . Ann. hist. nat. Mus. hung., Budapest, 74: 290, f 48. By courtesy. .

Gen. MARUSYLLUS gen. n. (21 species)
[name unpublished, proposed as partial synonym for the genus Yllenus (in part - hamifer group Prˇszynski, 1968)].

Type species Yllenus (hamifer group) hamifer Simon, 1876
ETYMOLOGY. Named in honor of prominent arachnologist - Dr. Y.M. Marusik [MARUS-ik-YLL-en-US], co-author of the big monograph of the genus Yllenus. Grammar gender - male.
DESCRIPTIONS. Cymbium striking by development of long, anterior "tail" , bending over palpal organ, with narrow depression along its whole length, guiding embolus, which is hair thin and very long. Bulbus relatively small, with conductor crescent shaped, broad and robust. Palpal femur broad, crescent shaped, tibia short with large tibial apophysis, articulating with corresponding plate-like outgrowth of the cymbium's postero-vental angle. Spermathecae relatively small and simple, copulatory ducts thin, varying complicated coils to very complicates double spiral. Length of ducts seem to parallel length of embolus.

Yllenus (hamifer group) hamifer :Proszynski J. 1968e. Ann. zool. 26: 430-434, f 2, 17, 32, 45, 57, 78-84. By courtesy.
Yllenus (hamifer group) auspex (a+b) + c) bator :Proszynski J. 1968e. Ann. zool. 26: 445-450, f 108-110
+c) 444-445, f. 46, 59, 98-100 +b) Logunovet al. 2011 .Arthr. Sel. 20: 239, f 16-18. By courtesy
Yllenus (hamifer group) maoniuensis +namulinensis + pamiricus: Logunov, Marusik 2003b: 151, f 493-496 + 136,
f 497-498 + 146-149, f 505-511. By courtesy.
Yllenus gregoryi +b) karnai : Logunov 2010. Bull. Br. arachnol. Soc.: 15 (3), 91–92, figs 1-2.4 +b) Logunov, Marusik 2003b: 138-139, f 466-471. By courtesy.

Gen. Yllenus Simon,1876
expected but not yet reported from Indian Subcontinent

[syn. Yllenus (arenarius group) Prószynski,1968]

Type species Yllenus arenarius Simon, 1876.
DIAGNOSIS. Male palps with robust cymbium and bulbus, conductor very robust, tibial apophysis prominent to enormous, articulating with corresponding structures of cymbium's wall. Epigyne with median pocket developed at the posterior edge, often bending ventral wards. Copulatory ducts thick walled, usually short and gently bent, spermathecae thick walled.

Yllenus (arenarius group) arenarius :Proszynski J. 1968e. Annales zoologici, 26: 488-492, f 9, 29, 44, 56, 71, 178-184 + Logunov, Marusik 2003b. KMK Sci.Press: 85-89, f 56 + ©Photo M. Bartos, 2000. By courtesy.
Yllenus (arenarius group) horvathi :Proszynski J. 1968e. Ann. zool. 26: 481-488, f 10, 30, 43, 55, 70, 170-177
+ © Photo: T. Szuts. By courtesy.