RETURN TO DATABASE - ALTERNATIVE CLASSIFICATION
SALTICIDAE OF THE WORLD - 0-INDEX of Genera - 01-FOREWORD & KEY to groups of genera! - 02-Comparison - HISTORICAL classification of Aelurilleae - 03-AEURILLINES - 04-AMYCINES- 05-AMYCOIDA VARIA - 06-ASTIAINES - BALLINES - 07-BELIPPINES - 08-CHRYSILLINES - 09-COCALODINES - 10-COLONINES[=Thiodinines] - 11-DENDRYPHANTINES - 12-DIOLENINES - EUODENINES - 13-EUOPHRYINES-PART 0: Introduction-Diversity - 13-EUOPHRYINES-PART 1: Admestina-Donoessus - 14-EUOPHRYINES-PART 2: Echeclus-Pystira - 15-EUOPHRYINES-PART 3: Rhyphelia-Zenodorus - 16-EUPOAINES - 17-EVARCHINES - 18-FOSSILS - 19-HABRONATTINES - 20-HARMOCHIRINES - 21-HELIOPHANINES - 22-HISPONINES - 23-HYLLINES - 24-ICIINES - 25-LAPSIINES - 26-LIGONIPEINES - 27-LYSSOMANINES - 28-MENEMERINES - 29-MYRMARACHNINES - 30-NOTICIINES - 31-PELLENINES - 32-PSEUDICIINES - 33-SIMAETHINES - 34-SITTICINES - 35-SPARTAEINES - 35a-SYNAGELINES - 36-THIRATOSCIRTINAE - 37-YAGINUMAELLINES - 38-YLLENINES - 39-UNCLASSIFIED TEMPORARY
APPENDICES -AP1-Simon's classification - AP2-Petrunkevich synthesis-1928 - AP3-Bonnet's list of subfamilies - AP4-Prószynski's revision of subfamilies 1976 - AP5-Prószynski - summary of results - AP6-Aelurillus black & gray - AP7-Maddison's views on Salticidae phylogeny 2014 - MS1-Omoedus synonymy
MADDISON'S ALTERNATIVE: Tittle_pg Index Introduction Agoriini Aelurillina Asemoneinae Amycini Amycoida-incertae sedis Astiini Ballini Baviini Bredini Chrysillini Cocalodini Dendryphantina Euophryini-1 Euophryini-2 Euophryini-3 Eupoinae Freyina Gophoini Harmochirina Hasariini Hisponinae Holcolaetina Huriini Itatina Lapsiini Leptorchestini Lyssomaninae Marpissina Mopsini Myrmarachnini Nannenini Neonini Onomastinae Plexippina Salticini Sarindini Scopocirini Simaethina Simonellini Sitticini Spartaeina Synagelina Thiodinini Thiratoscirtina Tisanibini Viciriini Salticidae-incertae sedis

Monograph of the Salticidae (Araneae) of the World 1995-2015. Part I: Introduction to alternative classification of Salticidae.

Jerzy Prószyński
36 - THIRATOSCIRTINES GROUP OF GENERA
sensu Bodner & Maddison 2012 - pending revision

Version July 1st, 2016.
Sources and permissions of illustrations, also references, are displayed in the IInd part of the monograph http://www.peckhamia.com/salticidae/

Interactive index of genera. Type genus Thiratoscirtus
Ajaraneola Alfenus Ansienulina Bacelarella "Bacelarella" Gramenca Lamottella Longarenus Malloneta Massagris "Massagris" schisma Nimbarus Pochyta Saraina Thiratoscirtus Ugandinella.
(52 species)

Introduction

Bodner & Maddison (2012, p. 221-222) proposed a new clade "THIRATOSCIRTINAE", which they consider subfamily, and discuss its evolutionary significance. Understanding premises for that is difficult because their taxonomic documentation is limited to listing genera they propose to include into THIRATOSCIRTINAE, without taxonomic definition or description, apart from a cladogram based on molecular analyses, whose taxonomic significance is unclear. To provide some orientation in their proposal I quote below the taxonomic documentation of all species they consider.

Remarks of relationships between genera of Thiratoscirtinae and other clades of Salticidae. Morphological documentation available in the literature is insufficient and therefore inconclusive. Structure of epigyne and palps represent two types, which deserve further attention, there are also some rather incompatible species. The most tempting similarity is separation of eyes anterior lateral and posterior lateral by prominent depression, with microscopic size of eyes II, supposedly indicating relationships to fossil Eocene Prolinus, and its extant analogues Tomocyrba and Hispo, looks probable in the case of Massagiris schisma, (whose relationship to Thiratoscirtinae is, however, questionable), but is not supported by eyes of Thiratoscirtus and related genera. Coiled embolus may indicate relationships to Euophryinae, but require further documentation of its basis. There is intriguing resemblance in posterior part of epigyne with Asiatic genus Habrocestoides, with curious circular structure inside, but that require further study. In view of very incomplete morphological documentation of diversity in Thiratoscirtinae, no firm conclusion on their placement could be drawn.
Good illustration of Bodner and Maddison views is the plate of 24 unidentified species - since I have no current copyright permission to display the plate, I direct readers to the original publication, including here thumbnails advertisement for that (below).
Advertisement for Bodner, Maddison 2012: 65: 237-238, plate B1-2 with 24 unidentified Thiratoscirtinae [?].
Remarks on genus Thiratoscirtus and related forms was given by Wesołowska 2011 (below). The documentation of Thiratoscirtus patagonicus, the type species of the genus Thiratoscirtus, prepared by Szuts and documenting its African relationships, has been reproduced in the database of Salticidae by Prószynski, 2003b (by kind permission of T. Szuts).
Definition by Wesolowska, Russel-Smith 2011. Ann. zool. 61(3): 603-604, f 188-192, 252. By courtesy.

Pictorial guide to identification of genera

Comparison of selected characters

Gen. Ajaraneola Wesołowska, Russel-Smith, 2011 (1 species)

Type species Ajaraneola mastigophora.
Diagnostic illustrations below are integral part of the definition.

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Ajaraneola mastigophora : Wesolowska, Russel-Smith.Ann. zool. 61(3): 558-559, f 14-20. By courtesy.

Gen. Alfenus Simon, 1902 (2 species)

Type species Alfenus calamistratus .
Diagnostic illustrations below are integral part of the definition.

Alfenus calamistratus : Szuts T., Scharff N. 2005. Acta zool. Acad. Sci. Hungar. 51: 359-361, f 1a,c, 2a-f, 3a-d. By courtesy.
Alfenus chrysophaeus : Szuts T., Scharff N. 2005. Acta zool. Acad. Sci. Hungar. 51: 361, f 1a,c, 2a-f, 3a-d.. By courtesy.

Gen. Ansienulina Wesołowska, 2015 (1 species)

Type species Ansienulina mirabilis .
Diagnostic illustrations below are integral part of the definition.

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Ansienulina mirabilis: Wesolowska, 2015: 56(2): 478, f. 1-14. By courtesy.. By courtesy.

Gen. Bacelarella Berland, Millot, 1941 (4 species)

Type species Bacelarella fradei .
Diagnostic illustrations below are integral part of the definition.

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+
Bacelarella fradei : Proszynski 1987. Atlas: 4. [Syntypes] + Rollard, Wesolowska. 2002. Zoosystema. 24 (2): 286, figs 2A-E
+ Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 88-91, figs 3F, 3M, 5B, 5E, 7B, 7E, 8C, 8F, 9D. By courtesy.
+
Bacelarella dracula: Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 86-88, figs 1B, 2A, 3D, 3K, 4C, 4D,
4G, 4H, 7C, 7F, 8B, 9B + Wesołowska, Russel-Smith 2011. Ann. zool. 61(3): 559-561, f 217-218. By courtesy.
Bacelarella iactans : Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 90-91, figs 3E, 3L,5A, 5D,6, 8C, 9C..
Bacelarella tanohi : Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 91-92, figs 1A, 3G, 3N, 5C, 5F, 8E.. By courtesy.

Gen. "Bacelarella" (4 species)
different genus, to be described as new

Type species Bacelarella ? .
Diagnostic illustrations below are integral part of the definition.

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"Bacelarella" conjugans : Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 83-85, figs 1C-D, 2B-E, 3C, J, 4B, F, 7A, D, 8A, 9A. By courtesy.
b)
"Bacelarella" pavida +b) tentativa: Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 83-85,
figs 3B, 3I, 4A, 4E +b) 285: 82-83, figs 3A, H . By courtesy.
"Bacelarella-2" gibbosa : [third genus?] Wesołowska, Edwards 2012. Ann. zool.: 62(4): 735, f 5–11, 111–113. By courtesy.

Gen. Gramenca Rollard, Wesołowska, 2002 (1 species)

Type species Gramenca prima.
Diagnostic illustrations below are integral part of the definition.

Gramenca prima : Rollard Ch., Wesolowska W. 2002.
Zoosystema. Paris, 24 (2): 292-293, figs 7A-D. By courtesy.

Gen. Lamottella Rollard, Wesołowska, 2002 (1 species)

Type species Lamottella longipes.
Diagnostic illustrations below are integral part of the definition.

Lamottella longipes : Rollard Ch., Wesolowska W. 2002. Zoosystema. 24 (2): 296, figs 10A-F. By courtesy.

Gen. Longarenus Simon, 1903 (1 species)

Type species Longarenus brachycephalus.
Diagnostic illustrations below are integral part of the definition.

Longarenus sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 122. By courtesy.
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Longarenus brachycephalus : Szuts T. 2005 [2007]. Opusc. zool., 36: 89-90, f 16-21 + Wesolowska, Edwards 2012.
Ann. zool.: 62(4): 748, f 51–59, 124. By courtesy.

Gen. Malloneta Peckham, Peckham, 1901 (1 species)

Type species Malloneta guineensis .
Diagnostic illustrations below are integral part of the definition.

Malloneta sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 124. By courtesy.
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Malloneta guineensis : Proszynski J. 1987. Atlas ...: 62. [Lectotype, Freetown [Sierra Leone] + Wesolowska, Edwards 2012. Ann. zool.: 62(4): 750-751, f 60–62, 125. By courtesy.

Gen. Massagris Simon, 1900 (7 species)

Type species Massagris constricta.
Diagnostic illustrations below are integral part of the definition.

Massagris sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 26-27. By courtesy.
b)c)
Massagris constricta +b) natalensis +c) contortuplicata : Simon 190a. 451, 452 +b) Wesolowska, Haddad 2009. Afr. Inver. 50(1):
53,f 93-98, 218 +c) 208-210, f 89, 103–106. By courtesy.
b)
Massagris honesta +b) mirifica: Wesołowska & Haddad 2013. Afr. Invert.: 54 (1): 210-212, f 19, 90, 91, 107–113 +b) Wesolowska 1993c. Genus, 4 (2): 137, figs 1-8.. By courtesy.
Massagris mohale: Wesolowska W., Haddad C.R., 2014 . Lesotho. Afr. Invert. 55 (2), 250-253, f 69, 70, 74–81.. By courtesy.
b)
Massagris separata +b) schisma [misplaced?]: Wesolowska 1993c. Genus, 4 (2): 137-138, figs 9-14 +b) Maddison, Zhang J.
2006 . Zootaxa 1255: 32, illustrations 8-10, ©Magnolia Press [Note embolus similarity to Pochyta poissonii!]. By courtesy.

Gen. Nimbarus Rollard, Wesołowska, 2002 (1 species)

Type species Nimbarus pratensis.
Diagnostic illustrations below are integral part of the definition.

Nimbarus pratensis : Rollard Ch., Wesolowska W. 2002. Zoosystema. Paris, 24 (2): 301-303, f 14A-F. By courtesy.

Comparison of palps of 5 species, used by Maddison et al., 2008 for gene sequencing, with types of their supposed congeners.

Gen. Pochyta Simon, 1901 (9 species)

Type species Pochyta spinosa .
Diagnostic illustrations below are integral part of the definition.

b)c)d)e)
Pochyta spinosa +b) simoni +c) poissonii +d) perezi +e) remyi : Proszynski J. 1987. Atlas ...: 79 [Not a type. Is it correctly
identified? ] +b) Lessert R. 1925a. Rev. suisse zool., 31 (13): 505, f 87-91 +c) Berland, Millot 1941. 12, 2: 363, f 64 +d) 362 f 63
[note embolus similarity to Massagris schisma!] +e) 364, f 64. . By courtesy.
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Pochyta pulchra : Proszynski 1984c. Atlas ...: 70. ©Photo T. Szuts. By courtesy.
b)c)d)
Pochyta fastibilis +b) major +c) moschensis +d) ["pannosa" & 6 other MISIDENTIFIED] [true
"pannosa" is "species inquirenda"]
: Simon E. 1903a.: 722, f 867 A-B +b) 716, f. 863-864 (=K-L)
+c) Caporiacco 1947d. 40: 249, t 2, f 76 +d) Maddison, Bodner, Needham 2008. Zootaxa 1893: 52, f 5.
©Magnolia Press. By courtesy.

Gen. Saraina Wanless & Clark, 1975 (3 species)

Type species Saraina rubrofasciata.
Diagnostic illustrations below are integral part of the definition..

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Saraina rubrofasciata : Szuts T., Scharff N. 2005. Acta zool. Acad. Sci. Hungar. 51: 366-368, f 5a-g, 6a-c
+ Azarkina 2009: 293, FIG. 1. By courtesy.
+
Saraina deltshevi +b) kindamba: Azarkina 2009: 293-295, figs 2-8 +b) 295-296, figs 9-18. By courtesy.

Gen. Thiratoscirtus Simon, 1886 (17 species)

Type species Thiratoscirtus patagonicus.
Diagnostic illustrations below are integral part of the definition. Species name misleading, type specimen from Congo, not Patagonia!

Thiratoscirtus [?] sp. : Maddison 2015. Journal of Arachnology. 43: 231&#8211;292, f. 123. By courtesy.
>+
Thiratoscirtus patagonicus : Galiano 1963b. Physis, 23 (66): 457-458, t. 39, f 11-14 + ©Photo Szuts [type specimen + its label
+ new African specimens]. By courtesy.
b)
Thiratoscirtus rudyi [undescribed] +b) procerus : ©Photo Szuts [from Congo (Brazzaville): Sibiti]
+b) Wesołowska, Edwards 2012. Ann. zool.: 62(4): 761-762, f 100–101, 130. By courtesy.
b)c) d)
Thiratoscirtus atakpa +b) efik [misplaced] +c) torquatus +d) alveolus : Wesołowska, Edwards 2012. Ann. zool.: 62(4): 759-760,
f 95–96 +b) 760, f 97–99 +c) 762-763, f 102–107, 131 + d)Wesołowska Russel-Smith, 2011. Ann. zool. 61(3): 599-600, f 167-169.
b)
Thiratoscirtus bipaniculus +b) mirabilis : Wesołowska Russel-Smith 2011. Ann. zool. 61(3): 600, f 170-172, 236 +b) 603-604, f 188-192, 252. By courtesy.
b)
Thiratoscirtus gambari +b) monstrum : Wesołowska Russel-Smith, 2011. Ann. zool. 61(3): 601, f 173-184, 250, 251 +b) 601-602, f 193-195, 253. By courtesy.
b)
Thiratoscirtus obudu [misplaced] +b) vilis : Wesołowska Russel-Smith 2011. Ann. zool. 61(3): 606, 196-198, 254, 255
+b) 601-602, f 199, 200, 256. By courtesy.
b)
Thiratoscirtus harpago +b) yorubanus : Wesołowska Russel-Smith 2011. Ann. zool. 61(3): 601-602, f 185-187 +b) 608-609,
201-204. By courtesy.
Thiratoscirtus mastigophorus :Wiśniewski, Wesołowska 2013. ©Genus 24(2): 252-260, f 1-13. By courtesy.
Thiratoscirtus perspicuus :Wiśniewski, Wesołowska 2013. ©Genus 24(2): 256-260, f 14-23.. By courtesy.

Gen. Ugandinella Wesolowska, 2006 ( 1 species)

Type species Ugandinella formicula.
Diagnostic drawings below are integral part of definition.
REMARKS. Placement doubtfull due to incompatible structures of epigyne.

Ugandinella formicula : Wesolowska 2006. Ann. Zool. 56(2): 435-439, f 1-17. [PLACEMENT?]. By courtesy.