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OUT OF DATE CLASSIFICATORY SYSTEMS -Simon's classification - Petrunkevich synthesis-1928 - Bonnet's list of subfamilies - Prˇszynski's revision of subfamilies 1976.
Monograph of the Salticidae (Araneae) of the World 1995-2014
Introduction to alternative classification of Salticidae
a subchapter

A Key to Salticidae of the Indian Subcontinent
group of genera
SPARTAEINAE
trial version

by Jerzy Prószyński & John T.D. Caleb
Version June 7th, 2015

Disclaimer. This is a work in progress, continually being developed and revised. Proposed new taxonomic names, included here are not valid unless subsequently published in accepted taxonomic papers. Note on images. This introduction to a classification includes only species defined by diagnostic illustrations in the taxonomic literature and Internet, arranged by similarities of embolus and spermophor. Images are displayed as thumbnails to save space and allow for rapid scanning of supposedly closely-related species. Thumbnails are small versions of illustrations in other parts of the monograph, where are displayed together with particulars of their sources and permissions.

Abstract

Interactive index of genera. Type genus Spartaeus.
Brettus Cocalus Cyrba Gelotia Holcolaetis Meata Meleon Mintonia Neobrettus Paracyrba Phaeacius Portia Spartaeus Taraxella Veissela Wanlessia Yaginumanis.

Introduction

REMARKS. Group of genera SPARTAEINAE is exceptionally well studied, owing to series of papers by Wanless 1978f, 1979b, 1980a, 1981b, 1981c, 1984a, 1984b, 1985, 1987. A concise summary of all research, including recent studies by molecular methods, is given by Maddison et al. 2014. In this situation the present paper is limited to scan of diversity of palps and epigyne of all species.

SPARTAEINAE are recognizable by large, flattish bulbus, circular or oval, covered entirely by tegulum, with translucent spermophor following the contour. Embolus prominent, arising anteriorly on side of bulbus, encircling its anterior part, seldom longer than half of bulbus length. In females spermatheca form of a large chamber, globular or oval, copulatory ducts relatively simple. Genus Holcolaetis departs from that general type, so do a few genera more - see drawings below.
TRANSFERS OF GENERA. Sonoita has an additional sclerite on bulbus, independent from the basis of embolus, with a small but distinct sclerotized hook. That resembles COCALODINAE and I transfer the genus there. Tentatively I transfer Sparbambus there. To SPARTAEINAE I includ Thrandina. Genus Paracyrba is synonymised with Cyrba, its only species Paracyrba wanlessi being sister species to Cyrba ocellata, which it resembles closely by color pattern and genital structures.

Guide to identification of genera by male palps

Guide to identification of genera by internal structure of epigyne

Guide to identification of genera by epigyne

Guide to identification of genera by color pattern

Gen. BrettusThorell, 1895

Type species Brettus cingulatus .

Brettus cingulatus : Wanless 1979b. Bull. brit. Mus. nat. Hist. (Zool.): 185-187, f. 1A, C, E, G; 2A-B . By courtesy.
b)+
Brettus albolimbatus +b) cf. albolimbatus : Wanless 1979b. Bull. brit. Mus. nat. Hist. (Zool.): 188, f. 2C; 3C-D; 4A-B
+b) ©Photo Anurag + ©Photo Vipin Baliga. By courtesy.
+
Brettus anchorum : Jastrzebski 1997. Genus, 8 (3-4): 702-703, figs. 1-9 + Wanless 1979b. Bull. brit. Mus. nat. Hist.
(Zool.): 188, ff. 2F; 3B; 4C-D. By courtesy.

Gen. Cocalus Koch C.L., 1846

Type species Cocalus concolor.

b)+
Cocalus concolor +b) gibbosus : Proszynski 1984c: 158 ( Holotype) +b) Wanless 1981c. Bull. brit. Mus. nat. Hist. (Zool.),
41 (5): 256, f. 2A-F + Davies Todd, Zabka 1989. Mem. Queens. Mus., 27 (2): 196, t 4 + ęPhoto R. Whyte. By courtesy.
+b)
Cocalus limbatus + b) murinus : Wanless 1981c. Bull. brit. Mus. nat. Hist. (Zool.), 41 (5): 258-260, ff. 5A-D
+b) 256-258, ff. 3A-E. +:© Photo H. K. Tang. By courtesy.
Cocalus sp [nadu] : India: Tamil Nadu. ©Photo J. Caleb. By courtesy.

Gen. Cyrba Simon, 1876

Type species Cyrba algerina.

+
+
Cyrba algerina : Proszynski 2003a. Annales zoologici. 53 (1): 46-47, figs 153-157 + ©Photo J. Lissner (from Spain)
+ ©Photo B. Knoflach (from Greece) [conspecific?]. By courtesy.
+
Cyrba ocellata : Proszynski 1978a: 16, ff. 19, 21-24 + Davies Todd, Zabka 1989. Mem. Queens. Mus., 27 (2): 194, 198, t. 6
+ ©Photo J. Caleb (Tamil Nadu, in preparation for print, specimen alive + discolored by preservation). By courtesy.
Cyrba wanlessi [=Paracyrba w. - comb.n.] : Zabka, Kovac 1996. Senck. biol. 76(1/2): 154-156, f 1-19. By courtesy.

Gen. Gelotia Thorell, 1890

Type species Gelotia frenata .

b)c)
Gelotia frenata b) bimaculata c) bouchardi : Proszynski 1968a.77: 11-16, ff. 1, 3, 5, 7, 9 +b) 16-20, ff. 2, 4, 6, 8
+c) Prószyński, Deeleman-Reinhold, 2012. Arthr. select. 21(1): 38, 47-58. By courtesy.
Gelotia sp: ©Photo R R. Jackson. By courtesy.
b)
Gelotia lanka +b) robusta : Wijesinghe D.P. 1991b. J. NY Entom. Soc., New York, 99 (2): 274-277, figs 1-6
+b) Wanless 1984a. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 174-176, ff. 16C, 19A-F. By courtesy.

Gen. Holcolaetis Simon, 1886
expected but not yet reported from Indian Subcontinent

Type species Holcolaetis xerampelina .

>
Holcolaetis xerampelina : Wanless 1985. Bull. brit. Mus. nat. Hist. (Zool.), 48 (4): 254, f. 2A-J, 3A-C, 16A
+ Wesolowska & Cumming, 2011: J. Afrotrop. Zool. .7:79, f 11-12. By courtesy.
+
Holcolaetis zuluensis : Wanless 1985. Bull. brit. Mus. nat. Hist. (Zool.), London, 48 (4): 259, f 6A-G, 16C
+ Wesołowska, Haddad 2009. African Invertebrates, 50(1): 47-49, f 83-86.. By courtesy.
Holcolaetis vellerea : Wanless 1985. Bull. brit. Mus. nat. Hist. (Zool.), London, 48 (4): 255-256, ff. 1A-B, 4A-J, 5A-D,
12A-B, 15A-B, 16B, 17-19. + Wesolowska., van Harten 2007. F. Arabia, 23: 221-223, f 89-94 + Wesolowska,
Cumming 2008. Ann. zool. 58: 188, f 67-70 + ©Photo John and Frances Murphy. By courtesy.

Gen. Meata Żabka, 1985
expected but not yet reported from Indian Subcontinent

Type species Meata typica .

b)c)
Meata typica +b) zabkai +c) [ANALOGY ONLY [?]: Brettus celebensis : Zabka 1985. Ann. zool. 39, 11: 239, ff. 279-282.
Prószyński, Deeleman-Reinhold, 2010. Arthr. selecta, 19(3): 174, f 107-110 - compared with M. typica. By courtesy.

Gen. Meleon Wanless, 1984
expected but not yet reported from Indian Subcontinent

Type species Meleon kenti.

Meleon kenti : Wanless F.R. 1978f. Bull. brit. Mus. nat. Hist. (Zool.)., London, 34 (3): 90-91, ff. 2 a-d +34(3): 92, f 3F.
b)
Meleon madagascarensis +b) raharizonina : Azarkina, Logunov 2010. Afr. Invert. 50(1): 173-176, f 40, 41, 42-47, 53-56
+b) 177, f 37, 48-51. By courtesy.

Gen. Mintonia Wanless, 1984
expected but not yet reported from Indian Subcontinent

Type species Mintonia tauricornis.

+
Mintonia tauricornis : Prˇszynski, Deeleman-Reinhold, 2012. Arthr. select. 21(1): 43-45, f 87-95
+ Wanless 1984. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 158-160, ff. 8A-H, 31A, 32E-F, 35B.. By courtesy.
.
b)
Mintonia bani +b) sp [Australia] : Ikeda, H. 1995b. Acta. arachn. 44: 117-121, figs 1-6.
+b) Davies Todd, Zabka 1989. Mem. Queensland Mus., 27 (2): 197, t. 5. By courtesy.
b)
Mintonia sp [AUS-1256] +b) sp [Borneo] : Queensland. ©Photo R. Whyte and I.R. Macaulay +b) ©Photo
P. Koomen. By courtesy.

Gen. Neobrettus Wanless, 1984
expected but not yet reported from Indian Subcontinent

Type species Neobrettus tibialis .

++
Neobrettus tibialis : Proszynski 1978a. Ent. bas. 19-21, f 25-27 + Wanless 1984a. Bull. brit. Mus. nat. Hist. (Zool.),
46 (2): 183-185, f 24A-H + ęPhoto John Murphy, 1997. By courtesy.
>
Neobrettus sp [Philippines] : ęPhoto R.R. Jackson [Philippines: Mt. Banaham]. By courtesy.

Gen. Paracyrba Żabka, Kovac 1996 = Cyrba Simon, 1876 (syn. n.)
Paracyrba wanlessi = Cyrba wanlessi (comb. n.) .

Gen. Phaeacius Simon, 1900 (12 species)

Type species Phaeacius fimbriatus.

++b)
Phaeacius fimbriatus +) azarkinae : Wanless F.R. 1981b. Bull. brit. Mus. nat. Hist. (Zool.), 41 (4): 202-205, f 4A-H, 5B
+ Prˇszynski, Deeleman-Reinhold, 2012. Arthr. select.21(1): 55, f 130-140 + 2010: 19(3): 178, 127-128. By courtesy.
+
Phaeacius lancearius : Proszynski 1984c. Atlas ...: 99 Syntype + 19(3): 178, 127-128 + Wanless F.R. 1981b.
Bull. brit. Mus. nat. Hist. (Zool.), 41 (4): 205, f. 1, 2A-F, 5A, C-F. By courtesy.
++
++
Phaeacius malayensis : Zhang J., Li D. 2005. 53: 221-229 + .19(3): 178, 127-128 + Wanless 1981b. 41 (4): 205-208, f 6A-E;
7A-C..+ ęPhoto J. Koh 1989 + © Photo H.K. Tang + ©Photo Hirzi Hussain + ©Photo Murphy F. and J. 2000. By courtesy.
b)
Phaeacius sp [Kerala 1] +b) sp [Baliga] : © Photo Samson Davis (Kerala) +b). ©Photo Vipin Baliga (S-Kodagu District, Karnataka). By courtesy.

Gen. Portia Karsch, 1878

Type species Portia fimbriata.

>+
Portia fimbriata : Wanless 1978f. Bull. brit. Mus. nat. Hist. (Zool.). 99-100, f 7A-G; 8A-F; t 3a-f; 4c-f; 5c, d, f
+ Davies Todd V., Zabka M. 1989. Mem. Queens. 27 (2): 195, t 3 + ©Photo Vipin Baliga (India). By courtesy.
+
Portia labiata : ę Photo Koh J. + H.K. Tang. By courtesy.
++b)
Portia albimana +b) sp [Chak 30] : Wanless 1978f. Bull. brit. Mus. nat. Hist. (Zool.). 34 (3): 107-108, f 12A-D.
+ Zabka 1985. Ann. zool., 39, 11: 437, f 487-490 + Murphy, Murphy 1983a. Bull. Br. arachnol. Soc.: 40, f 4-5
+b) © Photo Dr. S. Chakrabarti (India). By courtesy.
Portia assamensis: Wanless 1978f. Bull. brit. Mus. nat. Hist. 105-107,
f 10 d-f, 11 d-f. By courtesy.

Gen. Spartaeus Thorell, 1891

Type species Spartaeus spinimanus.


>>
Spartaeus spinimanus : Prˇszynski, Deeleman-Reinhold, 2010. Arthr. selecta.19(3): 180, f 134-137 + Wanless 1984.
Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 148-151, f 3A-F, 4A-G, 30A-D, 33F, 35A +ęPhoto D. Petot. By courtesy

++
Spartaeus thailandicus : Prˇszynski Internet drawings: Khao Yai National Park, coll. Deeleman-Reinhold + Logunov, Azarkina
2007: Arthr. Sel. 16(2): 110, f 86-94 + Wanless 1984. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 151-152, f 5A-D. By courtesy.

Gen. Taraxella Wanless, 1984
expected but not yet reported from Indian Subcontinent

Type species Taraxella solitaria.

b)>
Taraxella solitaria +b) sumatrana : Wanless, 1987 Bull. British Mus. (NH) (Zool. ser.) 52(3): 156-157, f 7A-G
+b) 125-128, t 12 A-J + Prˇszynski, Deeleman-Reinhold, 2012. Arthr. select. 21(1): 57-58, f 154-158. By courtesy.

Gen. Veissella Wanless, 1984
expected but not yet reported from Indian Subcontinent

Type species Veissella durbani.

+b)
Veissella durbani +b) milloti :Wesolowska & Haddad, 2009: 95, f. 205-209 + Wanless 1984a. Bull. brit. Mus. nat. Hist.
(Zool.), 46 (2): 189, f. 27A-G +b) Logunov & Azarkina, 2007: 16(2): 113, f 107-113. By courtesy.

Gen. Wanlessia Wijesinghe, 1992
expected but not yet reported from Indian Subcontinent

Type species Wanlessia sedgwicki.

Wanlessia denticulata : Peng X., Tso I., Li S. 2002. Zoological Studies 41 (1): 5, f 17-20. By courtesy.

Gen. Yaginumanis Wanless, 1984
expected but not yet reported from Indian Subcontinent

Type species Yaginumanis sexdentatus.

>+
b)+
Yaginumanis sexdentatus +b) saganus [interpreted by Wanless as female of Y. sexdentatus] : Bohdanowicz, Proszynski
1987. Ann. zool. 41, 2: 46-47, f 1-5 + © Ono, Ikeda, Kono 2009: 562, pl 44-1+b) Bohdanowicz, Proszynski 1987.
Ann. zool. 41, 2: 46-47, f 6-7 + Wanless 1984. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 153-155, f 6g-i. By courtesy.