RETURN TO DATABASE - ALTERNATIVE CLASSIFICATION
SALTICIDAE OF THE WORLD - 0-INDEX of Genera - 01-FOREWORD & KEY to groups of genera! - 02-Comparison - HISTORICAL classification of Aelurilleae - 03-AEURILLINES - 04-AMYCINES- 05-AMYCOIDA VARIA - 06-ASTIAINES - BALLINES - 07-BELIPPINES - 08-CHRYSILLINES - 09-COCALODINES - 10-COLONINES[=Thiodinines] - 11-DENDRYPHANTINES - 12-DIOLENINES - EUODENINES - 13-EUOPHRYINES-PART 0: Introduction-Diversity - 13-EUOPHRYINES-PART 1: Admestina-Donoessus - 14-EUOPHRYINES-PART 2: Echeclus-Pystira - 15-EUOPHRYINES-PART 3: Rhyphelia-Zenodorus - 16-EUPOAINES - 17-EVARCHINES - 18-FOSSILS - 19-HABRONATTINES - 20-HARMOCHIRINES - 21-HELIOPHANINES - 22-HISPONINES - 23-HYLLINES - 24-ICIINES - 25-LAPSIINES - 26-LIGONIPEINES - 27-LYSSOMANINES - 28-MENEMERINES - 29-MYRMARACHNINES - 30-NOTICIINES - 31-PELLENINES - 32-PSEUDICIINES - 33-SIMAETHINES - 34-SITTICINES - 35-SPARTAEINES - 35a-SYNAGELINES - 36-THIRATOSCIRTINAE - 37-YAGINUMAELLINES - 38-YLLENINES - 39-UNCLASSIFIED TEMPORARY
APPENDICES -AP1-Simon's classification - AP2-Petrunkevich synthesis-1928 - AP3-Bonnet's list of subfamilies - AP4-Prˇszynski's revision of subfamilies 1976 - AP5-Prˇszynski - summary of results - AP6-Aelurillus black & gray - AP7-Maddison's views on Salticidae phylogeny 2014 - MS1-Omoedus synonymy

MADDISON'S ALTERNATIVE: Tittle_pg Index Introduction Agoriini Aelurillina Asemoneinae Amycini Amycoida-incertae sedis Astiini Ballini Baviini Bredini Chrysillini Cocalodini Dendryphantina Euophryini-1 Euophryini-2 Euophryini-3 Eupoinae Freyina Gophoini Harmochirina Hasariini Hisponinae Holcolaetina Huriini Itatina Lapsiini Leptorchestini Lyssomaninae Marpissina Mopsini Myrmarachnini Nannenini Neonini Onomastinae Plexippina Salticini Sarindini Scopocirini Simaethina Simonellini Sitticini Spartaeina Synagelina Thiodinini Thiratoscirtina Tisanibini Viciriini Salticidae-incertae sedis

Monograph of the Salticidae (Araneae) of the World 1995-2015. Part I: Introduction to alternative classification of Salticidae.

Comparison of classifications by Maddison 2015 and Prószyński 2016a
Compiled by Jerzy Prˇszyński

Maddison's alternative
M_17 - Thiodinini
Subfamily: Salticinae: Clade: Amycoida: Tribe: Thiodinini
Version July 1st, 2016.
Sources and permissions of illustrations are displayed in the IInd part of the monograph http://www.peckhamia.com/salticidae/

Type genus .
Index. Agelista Arachnomura Atomosphyrus Bredana Cyllodania Hyetussa Micalula
Thiodina Titanattus

Gen. Agelista Simon, 1900 (1 species)

Type species Agelista andina.

>
Agelista andina + sp. : Galiano 1963b.Physis, 23 (66): 283-285, t. 3, f 4-6
+ © Photo Machado & Gasnier - INPA023: Campus-UFAM-AM. By courtesy.

Gen. Arachnomura Mello-Leitao, 1917 ( 2 species)

Type species Arachnomura hieroglyphica.
DIAGNOSIS. Bulbus round, loop of spermophor antero-lateral, embolus encircles bulbus tightly. Cymbium broadened, with postero-lateral edge drawn, touching tibial apophysis. Tibia triangular. Epigyne with a pair of large, oval grooves. Copulatory ducts make several elongate coil parallel to body axis, producing thin sclerotized, lateral ducts, running to small, globular spermathecae anteriorly. Body average looking, strikingly different from Amycus and Asaracus. Diagnostic drawings below are integral part of definition.

Arachnomura sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 37. By courtesy.
b)
Arachnomura hieroglyphica + adfectuosa : Galiano 1977a. J. Arachnology 3: 143, ff. 7-10, 18-19 + © Photo C.J. Grismado.

Gen. Atomosphyrus Simon,1902 ( 2 species)

Type species Atomosphyrus tristiculus
DIAGNOSIS. Bulbus round, encircled tightly by embolus. Spermathecae anterior, small and oval, atop of narrow and long, lateral ducts [remaining details of drawing illegible]. Diagnostic drawings below are integral part of definition.

b)
Atomosphyrus tristiculus + breyeri: Galiano M. E. 1966a. Physis 26 (72): 282-284, ff. 9-16, 19, 20 + 280-281, f. 1-8 +b) Richardson, 2010. Zootaxa 2418: 1-49. By courtesy.

Gen. Bredana Gertsch, 1936 (1 species)

Type species Bredana complicata.
Diagnostic drawings below are integral part of definition.

Bredana complicata Gertsch W. J. 1936. Americ. Mus. Novitates, 852: 21, f 33-35. By courtesy.

Gen. Cyllodania Simon,1902 ( 2 species)

Type species Cyllodania bicruciata .
DIAGNOSIS. Embolus broad, with fleshy component, arises from broader basis, encircles entirely bulbus. Epigyne with large oval depression delimited incompletely by extending rims of copulatory openings, the latter extend transversally in posterior part of epigyne and lead to broad, transverse copulatory ducts, these pass through a few half-coils, narrowing abruptly and pass to duct shaped, narrow, slightly broadening spermatheca, running along axis and terminated by small, oval chamber in anterior part of epigyne. Diagnostic drawings below are integral part of definition.

Cyllodania sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 35. By courtesy.
Cyllodania bicruciata :Galiano 1977a. J. Arachn. 3: 138, f 11-12, 20-22, 27-28. By courtesy.
Cyllodania minuta :Galiano 1977c. Rev. per. Ent.: 20 (1): 77, f. 7-10, 13-17. By courtesy.

Gen. Hyetussa Simon, 1902 (6 species)

Type species Hyetussa simoni.
DIAGNOSE. Epigyne with a pair of large, membranous white windows, internally with beautiful spiral of narrow copulatory ducts, spermathecae are small, oval chambers in front of spiral, in the anterior part of epigyne. Embolus broad, encircles bulbus. Diagnostic drawings below are integral part of definition.

+
Hyetussa simoni + mesopotamica: Galiano 1976b. Physis C., 35(90): 64, figs 8-13 + 233-237, f 18-20, 27-30, 39, 45-50.
Hyetussa andalgalaensis :Galiano 1976b. Physis C., 35(90): 231, f. 21-24, 31-32, 36, 51 + ©Photo C.J.Grismado. By courtesy.
b)+
Hyetussa aguilari +b) cribrata: Galiano 1978a. Rev. Ent. 21 (1): 29, f. 8-14 +b) 2, t 2I, f. 6-10 + Galiano 1976b. 35 (91): 233, f 40-42.
Hyetussa secta : Galiano M. E. 1976b. Physis C., 35 (91): 239-241, ff. 25-26, 33-34. By courtesy.

Gen. Micalula (Petrunkevitch, 1925)

Type species Mica longithorax Petrunkevitch, 1925a: 246.

Micalula longithorax Petrunkevitch A. 1925, 27: 81, 246, f. 155-157.

Gen. Thiodina Simon, 1900 (3 species)
PLACEMENT PROVISIONAL

Type species Thiodina nicoleti
Thiodina nicoleti was recently revised by Bustamante, Maddison, Ruiz 2015 (Zootaxa 4012 (1): 181–190), who redefined the species and produced drawings of its palpal organ, which seem to place genus within group HYLLINES, characterized by embolus originating from posterior lateral end of bulbus, entirely different from COLONINES, as proposed in the present work. Female of Thiodina nicoleti remains unknown, so we cannot use its structure of spermathecae and ducts in classification. However, bulbous sensory setae ventrally on tibia I and on faces of chelicerae, according to Bustamante et al., 2015 diagnostic for the species, were not tested for occurrence in other genera, so are not sufficient character for classification of genera. Bustamante mentions as possible relative genus Hyetussa (AMYCINES), which has beautiful spiral of copulatory ducts in epigyne, its embolus has similar origin but is much longer, twisted around bulbus. With insufficient data on palps and internal structures of epigyne we will have to wait until further studies, promised by these authors.

b)
Thiodina nicoleti + neotype : Original by Nicolet - Richardson, 2010. Zootaxa 2418: 42, f 117-118 +b) Bustamante, Maddison, Ruiz 2015. Zootaxa 4012 (1): 181–190, f 1-30. © Magnolia Press. By courtesy.

Gen. Titanattus Peckham, Peckham, 1885 (6 species

Type species Titanattus saevus.
Diagnostic drawings below are integral part of definition.
REMARKS. Placement doubtfull due to cheliceral dentition, containing rows of teeth on two separate edges of chelicerae.

Titanattus sp. : Maddison 2015. Journal of Arachnology. 43: 231–292, f. 38. By courtesy.
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Titanattus saevus + pegaseus : F. P.-Cambridge 1900. 191, t 15, f 1.
+ Simon E. 1901a: 419, figs L-M (= 464-478) + Galiano 1963b. Physis, 23(66): 460, t 37, f 16-17 [PLACEMENT?]. By courtesy.
++
Titanattus cretatus + paganus + notabilis : Chickering 1946. Bull.Mus. Comp. Zool. Harvard 97: 383, f 337-340 + 385 f 341-342. + Galiano 1963b. Physis, 23(66):32, t 1, f 11; t 2, f 1-3. By courtesy.
Titanattus pallidus Edwards, Rinaldi, Ruiz 2005. Biota Neotr. 5(2): 20, f 1-3, 44-46.1-31. By courtesy.