RETURN TO DATABASE - ALTERNATIVE CLASSIFICATION
SALTICIDAE OF THE WORLD - 0-INDEX of Genera - 01-FOREWORD & KEY to groups of genera! - 02-Comparison - HISTORICAL classification of Aelurilleae - 03-AEURILLINES - 04-AMYCINES- 05-AMYCOIDA VARIA - 06-ASTIAINES - BALLINES - 07-BELIPPINES - 08-CHRYSILLINES - 09-COCALODINES - 10-COLONINES[=Thiodinines] - 11-DENDRYPHANTINES - 12-DIOLENINES - EUODENINES - 13-EUOPHRYINES-PART 0: Introduction-Diversity - 13-EUOPHRYINES-PART 1: Admestina-Donoessus - 14-EUOPHRYINES-PART 2: Echeclus-Pystira - 15-EUOPHRYINES-PART 3: Rhyphelia-Zenodorus - 16-EUPOAINES - 17-EVARCHINES - 18-FOSSILS - 19-HABRONATTINES - 20-HARMOCHIRINES - 21-HELIOPHANINES - 22-HISPONINES - 23-HYLLINES - 24-ICIINES - 25-LAPSIINES - 26-LIGONIPEINES - 27-LYSSOMANINES - 28-MENEMERINES - 29-MYRMARACHNINES - 30-NOTICIINES - 31-PELLENINES - 32-PSEUDICIINES - 33-SIMAETHINES - 34-SITTICINES - 35-SPARTAEINES - 35a-SYNAGELINES - 36-THIRATOSCIRTINAE - 37-YAGINUMAELLINES - 38-YLLENINES - 39-UNCLASSIFIED TEMPORARY
APPENDICES -AP1-Simon's classification - AP2-Petrunkevich synthesis-1928 - AP3-Bonnet's list of subfamilies - AP4-Prˇszynski's revision of subfamilies 1976 - AP5-Prˇszynski - summary of results - AP6-Aelurillus black & gray - AP7-Maddison's views on Salticidae phylogeny 2014 - MS1-Omoedus synonymy

MADDISON'S ALTERNATIVE: Tittle_pg Index Introduction Agoriini Aelurillina Asemoneinae Amycini Amycoida-incertae sedis Astiini Ballini Baviini Bredini Chrysillini Cocalodini Dendryphantina Euophryini-1 Euophryini-2 Euophryini-3 Eupoinae Freyina Gophoini Harmochirina Hasariini Hisponinae Holcolaetina Huriini Itatina Lapsiini Leptorchestini Lyssomaninae Marpissina Mopsini Myrmarachnini Nannenini Neonini Onomastinae Plexippina Salticini Sarindini Scopocirini Simaethina Simonellini Sitticini Spartaeina Synagelina Thiodinini Thiratoscirtina Tisanibini Viciriini Salticidae-incertae sedis

Monograph of the Salticidae (Araneae) of the World 1995-2015. Part I: Introduction to alternative classification of Salticidae.

Jerzy Prˇszyński
27 - LYSSOMANINES GROUP OF GENERA
Version July 1st, 2016.
Sources and permissions of illustrations, also references, are displayed in the IInd part of the monograph http://www.peckhamia.com/salticidae/

Type genus Lyssomanes
Interactive index of genera: Asemonea Chinoscopus Goleba Hindumanes Lyssomanes Macopaeus Onomastus Pandisus Sumakuru
.(129 species)

Introduction

LYSSOMANINES are defined by arrangement of eyes in four rows and by numerous separate teeth on inner posterior margin of chelicerae. Use of palp structures for identification require additional comparative research, internal structures of epigyne are simply unknown for majority of species. The genera may be provisionally separated by their external appearance. Drawings below are integral part of the definition.
REMARKS. Eyes arrangement in four rows occurs also in genera not related to LYSSOMANIDAE: Athamas and Leptathamas belongs to EUOPHRYINES, placement of Orthrus is unclear. For details of diagnostic characters - see comparison of species within particular genera. For different opinions see Maddison 2015.

Guide for identification of genera by general appearance

Gen. Asemonea Pickard-Cambridge O., 1869 (22 species)

Type species Asemonea tenuipes. Drawings below are integral part of the definition.

>+ b) c)
Asemonea tenuipes +b) serrata +c) pallida :: Peckham, Peckham, Wheeler 1889: 243, pl. 12, f. 5, 19 + Proszynski J. 1984c. Atlas: 3. + ęPhoto R.R.Jackson, 2000. +Wesolowska W. 2001b. Genus, 12(4): 581-582, f 11-14 + 582, figs 15-20. By courtesy.
b)
Asemonea amatola +b) cuprea :: Wesołowska & Haddad 2013. Afr. Invert.: 54 (1): 182-184, f 2, 30, 31 +b)Wesołowska, 2009. African zoology 44(1): 126-128, f 1-7 . By courtesy.
b)+ +
Asemonea flava +b) murphyae :: Wesolowska W. 2001b. Genus, 12(4): 578-579, figs 1-4 +b) 579-581, figs 5-10 + Wesołowska & Haddad 2013. Afr. Invert.: 54 (1): 185, f 34. + ęPhoto John and Frances Murphy. By courtesy.
b)c)
Asemonea virgea +b) crinita +c) liberiensis : Wesolowska W., Szuts T. 2003a.64: 59-62, figs 1-7 +b) Wanless1980d. Bull. brit. Mus. nat. Hist. (Zool.) 39 (4): 234, ff. 14A-E +c) 227-228, f. 8a-c. By courtesy.
b)c)
Asemonea cristata +b) fimbriata +c) maculata : Wanless1980d.Bull. brit. Mus. nat. Hist. (Zool.) 39 (4): 242, f. 19A-E +b) 229-230, ff. 10a-f +c) 233-234, f. 13a-e. By courtesy.
b)c) d)+
Asemonea minuta +b) ornatissima +c) picta +d) pinangensis:: Wanless1980d.Bull. brit. Mus. nat. Hist. (Zool.) 39 (4): 228-229, ff. 9a-e +b) 239-240, ff. 17A-D. +c) 243, f. 20A-C. +d) 244, ff. 21a-e. By courtesy.
b)c)d)
Asemonea pulchra+b) serrata +c) sichuanensis +d) trispila:: Wesołowska, Edwards 2012. Ann. zool.: 62(4): 734, f 1–4 +b) Wanless1980d.Bull. brit. Mus. nat. Hist. (Zool.) 39 (4): 234, f. 15A-H +c) Song, Zhu, Chen 1999: 505, figs 288 K-L +d) Tang G., Yin C., Peng X. 2006. Acta zootaxonomica sinica, 31: 547-548, f 1-3.. By courtesy.
+ +
b)
Asemonea stella [from Africa]+b) [from Australia] ::Wanless1980d.Bull. brit. Mus. nat. Hist. (Zool.) 39 (4): 237, f.16a-f + Wesolowska, Russel-Smith. 2000. Tropical Zool., 13 (1): 16-19 , figs 14-17+ Wesołowska, Haddad 2009. African Invertebrates, 50(1): 21, 14-15.+b) Szuts 2000. Folia entom. hungarica, LXI: 61-63, figs 1-6. By courtesy.
+
Asemonea tanikawai : Ikeda, 1996. Acta arachn., 45(2):113-117, figs 1-11 + Proszynski J. 1984c. Atlas: 3 + ©Photo Akio Tanikawa. By courtesy.
Asemonea clara : Wesołowska, Haddad, 2013: 54 (1): 184-185, f 32-33. By courtesy.

Gen. Chinoscopus Simon, 1901 (4 species)

Type species Chinoscopus gracilis. Drawings below are integral part of the definition.

Chinoscopus gracilis: Galiano 1998. Bull. Brit. Arachn. Soc. 11 (1): 5-7, f 26-27 + ęPhoto Gasnier & Machado.
b)
+c)
Chinoscopus ernsti +b) flavus +c) cf .flavus: Galiano 1998. Bull. Bri. Arach.. Soc. 11 (1): 7-8, fig. 14 +b) 7, figs 5-7 + ęPhoto A. Anker +c) Maddison 2015. Journal of Arachnology. 43: 231–292, f. 5. By courtesy. Note eyes AM color difference, serving as method of communication in Lyssomanidae, according to J. Crane .
++
Chinoscopus maculipes: Crane 1943: 128, ff. 1 l-o + Galiano 1998. Bull. Brit. Arachn. Soc. 11 (1): 8, figs 10, 15, 28, 29
+ Logunov, Marusik 2003c. Bull. Br. arachnol. Soc.,12 (9): 415, figs 1-10. By courtesy.

Gen. Goleba Wanless, 1980 (4 species)

Type species Goleba puella . Drawings below are integral part of the definition.

+
Goleba puella +b) sp. :Wanless 1980d. Bull. brit. Mus. nat. Hist. (Zool), 39 (4): 246-249, f. 22A-E; 23A-C; 29 + Wesolowska, Haddad 2009. African Invertebrates, 50(1): 38, f 53-61 + + ęPhoto R.R. Jackson, By courtesy.
Goleba jocquei : Szuts T. 2001. Annls Mus. R. Afr. Centr. (Zool.). 285: 111-116, figs 1-7. By courtesy.
>b)
Goleba pallens +b) punctata. : Wanless 1980d. Bull. brit. Mus. nat. Hist. (Zool), 39 (4): 249-250, ff. 24A-H + Saaristo M.I. 2010: 181, 27.37-42 +b) Wanless 1980d. Bull. brit. Mus. nat. Hist. (Zool), 39 (4): 249-250, ff. 24A-H. By courtesy.
Goleba lyra : Maddison, Zhang J. 2006 . Zootaxa 1255: 30, illustrations 1-7. . By courtesy.

Gen. Hindumanes Logunov 2004 (1 species)

Type species Hindumanes karnatakaensis . Drawings below are integral part of the definition.

Hindumanes karnatakaensis : Logunov D.V. 2004c. Bull. Br. arachnol. Soc. 13 (3): 73-75, f. 1-3. By courtesy.

Gen. Lyssomanes Hentz, 1845 (79 species)

Type species Lyssomanes viridis. Drawings below are integral part of the definition.

b)
c) >
Lyssomanes viridis [a + c] +b) jemineus :Galiano 1980b. Opera Lilloana 30: 45-46, f 101-103 +b) Galiano M.E. 1980b. Opera Lilloana 30: 63-65, t 20, f 150-152 +c) Photo D. Hill. By courtesy .
>
Lyssomanes longipes ::Galiano 1980b. Opera Lilloana 30: 12-14, f. 1-7 + ęPhoto Gasnier & Azevedo. By courtesy .
b)
Lyssomanes anchicaya +b) belgranoi :: Galiano M.E. 1984d. Bull. Br. arachnol. Soc. 6 (6): 268, 270, ff. 1, 2, 12-16
+b) 274, ff. 17-19, 31-32. By courtesy .
b)
Lyssomanes boraceia +b) blandus :: Galiano M.E. 1984d. Bull. Br. arachnol. Soc. 6 (6): 270, ff. 3, 35-36
+b)
Galiano 1980b. Opera Lilloana 30: 52, f. 129-13052, f. 129-130. By courtesy .
+
Lyssomanes camacanensis :: Galiano 1980b. Opera Lilloana 30: 42, f. 69-72, 77-79 + Logunov 2002. Reichenbachia, 34: 229-239, figs 1-2, 4-5.. By courtesy .
b)c)
Lyssomanes ceplaci +b) consimilis +c) diversus :: Galiano 1980b. Opera Lilloana 30 : 74-75, f. 167-174 +b) 34-36, f. 33-36 +c) 6 6, t. 19, ff. 145-148. By courtesy.
b)+
Lyssomanes devotoi +b) convexus :Galiano 1980b. Opera Lilloana 30: 37, f. 48 +b) 12-14, f. 1-7
+ Logunov, Marusik 2003c. Bull. Br. arachnol. Soc., 12 (9): 417-418, figs 13-14 . By courtesy .
b)c)
Lyssomanes deinognathus +b) dissimilis +c) ecuadoricus ::Galiano 1980b. Opera Lilloana 30: 69-71, t. 22, ff. 153-156 +b) 36-37, f. 46-47, 63-64 +c) Logunov, Marusik 2003c. Bull. Br. arachnol. Soc., 12 (9): 417-418, figs 20-21. By courtesy.
b)c)
Lyssomanes eatoni +b) elongatus +c) fossor :: Galiano 1980b. Opera Lilloana 30: 62-63, f. 121-12216, ff. 10-14
+b) 16, ff. 10-14 +c) Galiano M.E. 1996b. Iheringia, Ser. Zool. Porto Alegre, (81): 27-29, figs 11-13.
By courtesy .
b)
Lyssomanes hieroglyphicus +b) elegans ::Galiano 1980b. Opera Lilloana 30: 32-33, f. 27 +b) 47-48, t. 15 f. 117-118, t 16 ff 123-124. By courtesy .
b)
Lyssomanes ipanemae +b) miniaceus ::Galiano 1980b. Opera Lilloana 30: 48-49, ff. 110-116
+b) Galiano 1962d: 68-69, t. 5 ff. 6-9. . By courtesy .
b)
Lyssomanes jucari +b) janauari :: Galiano 1984d. Bull. Br. arachnol. Soc. 6 (6): 272, ff. 4, 33-34 +b) Logunov 2002.
Reichenbachia, 34: 229-239, figs 12-13. By courtesy .
b)c)
Lyssomanes lancetillae +b) leucomelas +c) minor:: Galiano 1980b. Opera Lill. 30: 38-39, ff. 50-51, 58-62+b) 22-23, ff. 52-54 +c) Galiano 1962d. Acta zool. Lill. 18: 69-70, t. 4, f. 8.. By courtesy .
b)
Lyssomanes limpidus +b) malinche :: Galiano M.E. 1984d. Bull. Br. arachnol. Soc. 6 (6): 274, f. 24-26 +b) Galiano 1980b. Opera Lilloana 30: 72-73, f. 179-182 . By courtesy .
+++
Lyssomanes mandibulatus :: Galiano 1980b. Opera Lilloana 30: 58-59, f. 92-95 + Logunov, Marusik 2003c.
Bull. Br. arachnol. Soc., 12 (9): 419-420, figs 22-28 . By courtesy .
+b)
Lyssomanes nigrofimbriatus +b) onkonensis : Galiano 1962d. Acta zool. Lill. 18: 70-73, t. 2, ff. 5-8 + Galiano 1980b. Opera Lilloana 30: 16-18, f. 15-16 +b) Logunov, Marusik 2003c. Bull. Br. arachnol. Soc., 12 (9): 420, figs 29-40. By courtesy .
b)
Lyssomanes nigropictus +b) peruensis: Galiano M. E. 1962d. Acta zool. Lill. 18: 73-74, t. 4 ff. 1-3 + Galiano 1980b. Opera Lilloana 30: 12-14, f. 1-7 +b) Logunov 2000a. Reichenbachia, 33: 267, figs 6-9. By courtesy .
+b)c)
Lyssomanes parallelus +b) parki +c) paravelox::Galiano 1962d. Acta zool. Lill. 18: 58, t 1, f 4-7 + Galiano 1980b. Opera Lilloana 30: 39-40, f. 49 +b) 51-52, f. 127-128 +c) Logunov 2002. Reichenbachia, 34: 232, figs 20-22. By courtesy .
b)c)d)
Lyssomanes patens +b) penicillatus+c) pichilingue d) placidus: Galiano 1980b. Opera Lilloana 30: 49-51, f. 107-109, 131-132 +b) 24-25, f. 56-57 +c) Galiano . 1984d. Bull. Br. arachnol. Soc. 6 (6): 270, ff. 5-9 +d) F. P.-Cambridge 1900. 180, 183, t. 13, f. 13. By courtesy .
b)c)d)
Lyssomanes portoricensis +b) protarsalis +c) reductus +d) robustus: Galiano 1980b. Opera Lilloana 30: 29-30, f. 23-26
+b) 59-60, ff. 87-89 +c) 60, f. 82-86 +d) 19-21, f. 38-40. By courtesy .
++
Lyssomanes quadrinotatus ::Galiano 1962d. Acta zool. Lill. 18: 85-86, t. 7 ff. 5-8 + Galiano 1980b. Opera Lilloana 30: 46-48,
ff. 105-106. By courtesy .
b
Lyssomanes remotus +b) santarem:: Galiano 1980b. Opera Lilloana 30: 54-56, f. 96-100 +b) 270, ff. 27-28 ęPhoto Gasnier. By courtesy .
b)c)d)
Lyssomanes [Fossil - 20-30 Myr old] +b) romani +c) trinidadus +d) waorani: Garcia-Villafuerte, Penney 2003. J. Arachn.
31: 400-404, f 1-2 +b) Logunov , Marusik 2003a. Bull. Br. arachnol. Soc., 12 (9): 420-422, figs 17-19 +c) 422-423, figs 11-12
+d) 422-424, figs 15-16. By courtesy .
+b)c)d)e)
Lyssomanes spiralis +b) sylvicola +c) tapuiramae +d) temperatus +e) tapirapensis : F. P.-Cambridge 1900.180, 183, t 12, f 11 + Galiano 1980b. Opera Lilloana 30: 65-66, t. 18, f. 138-139 +b) 73-74, f. 175-178 +c) 44, f. 73-76 +d) 67-68, t. 19, f. 141-144 +e) Galiano 1996b. Iheringia, Ser. (81): 26-27, figs 8-10. By courtesy .
b)
Lyssomanes tarmae +b) pescadero:: Galiano 1980b. Opera Lilloana 30: 71-72, ff. 161-166 +b) Jimenez, Tejas 1993. J. Arachnology, 207,f 9-15. By courtesy . .
b)+
Lyssomanes tristis +b) tenuis:: Galiano 1980b. Opera Lilloana 30: 40-42, f. 65-68, 80-81 +b) 14, f. 8-9 + Galiano 1962d. Acta zool. Lill. 18: 86, 88, t. 6 ff. 4-6. By courtesy .
b)+
Lyssomanes unicolor +b) velox::Galiano 1980b. Opera Lilloana 30: 56-58, t. 12, f. 90-91 +b) Galiano 1962d. Acta zool. Lill. 18: 90-92, t. 7, ff. 1-4 + Logunov, Marusik 2003c. Bull. Br. arachnol. Soc., 12 (9): 423-424, figs 46-50. By courtesy .
b)
Lyssomanes vinocurae +b) lehtineni : Galiano M.E. 1996b. Iheringia, Ser. Zool., (81): 24-26, figs 1-7 +b) Logunov 2000a. Reichenbachia, Dresden, 33: 268-269, figs 1-2 . By courtesy .
Lyssomanes yacui ::Galiano 1980b. Opera Lilloana 30: 12-14, f. 1-7 . By courtesy .
+b) +c)
Lyssomanes taczanowskii : Galiano 1980b. Opera Lilloana 30: 26-28, ff. 41-45 +b) W.P. Maddison et al. (2016) ZooKeys 549: 7, f. 8.+c) Logunov, Deza, 2010. A. arachn. 59(1): 21-23, f 1-3 [CONSPECIFIC?]. By courtesy .
+
Lyssomanes pauper ::Galiano 1980b. Opera Lilloana 30: 12-14, f. 1-7 + Logunov 2002. Reichenbachia, 34: 229-239, figs 1-2, 4-5. By courtesy .
b)
Lyssomanes flagellum +b) euriensis:: Kraus O. 1955b. 493: 57, figs 159-162. +b) Logunov, 2010. Acta arachn. 59(1): 21-23, f 1-3. By courtesy ..
b)
Lyssomanes burrera +b) austerus :: Jimenez, Tejas 1993. J. Arachnology, 205,f 1-8 +b) Galiano M.E. 1980b. Opera Lilloana 30: 21-22, f. 55 . By courtesy .
b)
Lyssomanes bryantae +c) blandus ::Galiano 1980b. Opera Lilloana 30: 53, f. 119-120 +b) 52, f. 129-130. By courtesy .
b)+
Lyssomanes bitaeniatus +b) benderi ::Galiano 1980b. Opera Lilloana 30: 68-69, f. 157-160 +b) Logunov 2002. Reichenbachia, Dresden, 34: 229-239, figs 8-13, 16-18 + Logunov & Marusik, 2003b: 415: [female described originally as L. pseudobenderi].
<b)<
+
Lyssomanes antillanus + amazonicus :: Galiano 1962d. Acta zool. Lill. 18: 52-53, t 3, f 7-9 + Galiano 1980b. Opera Lilloana 30:
28-29, f. 17-22 . +b) 33-34, f. 37 + Galiano 1962d. Acta zool. Lill. 18: 49-52, t 1, f 1-3 + ©Photo Gasnier. By courtesy .
Lyssomanes adisi :: Logunov 2002. Reichenbachia, 34: 229-239, f 4-5. . By courtesy .

Gen.Macopaeus Simon, 1900 (1 species)

Type species Macopaeus spinosus . Drawings below are integral part of the definition.

Macopaeus spinosus :Wanless F.R. 1980a. Bull. brit. Mus. nat. Hist. (Zool.)., 38 (4): 220, ff. 1A-F. By courtesy.

Gen. Onomastus Simon, 1900 (12 species)

Type species Onomastus nigricaudus . Drawings below are integral part of the definition.

Onomastus sp.: Maddison 2015. Journal of Arachnology. 43: 231&#8211;292, f. 2. By courtesy.
>
Onomastus nigricaudus :Wanless 1980c. Bull. brit. Mus. nat. Hist. (Zool.)., 39 (3): 181-183, f 1A-G. By courtesy.
Onomastus complexipalpis + O. danum + O. kaharian + O. simoni : Prˇszyński & Deeleman-Reinhold, 2013. Arthr selecta. 21(2): 134-135, f 89-107. By courtesy.
+
Onomastus kanoi : Ono H. 1995. Bull. National Science Museum, A (Zoology). 21 (3): 165, f 16-21 +©Photo Akio Tanikawa. By courtesy.
Onomastus - all species described in the monograph : Benjamin S.P. 2010 Revision and cladistic analysis of the jumping spider genus Onomastus (Araneae: Salticidae). Zoological Journal of the Linnean Society, London 159: 711–745, f  1-28. By courtesy.
>b)
Onomastus patellaris+b) quinquenotatus: Wanless 1980c. Bull. brit. Mus. nat. Hist. (Zool.)., 39 (3): 183-187, f 3A-G +b) 183, f 2A-E. By courtesy.

Gen. Pandisus Simon, 1900 (6 species)

Type species Pandisus scalaris.Drawings below are integral part of the definition.

b)c)
Pandisus scalaris +b) decorus +c) indicus:Wanless 1980d. Bull. brit. Mus. nat. Hist. (Zool.), 39 (4): 219, f. 3
+b) 221-222, f 5a-g +c) Proszynski 1992b. Ann. zool., 44, 9: 194, f 24, 27-28 . By courtesy.
b)c)
Pandisus modestus +b) parvulus +c) sarae: Wanless 1980d. Bull. brit. Mus. nat. Hist. (Zool.), 39 (4): 222-224, f 6A-D +b) 225, f 7A-E +c) 219-221, f 4a-j . By courtesy.

Gen. Sumakuru Maddison, 2016 (1 species)

Type species Sumakuru bigal Maddison, 2016. Drawings below are integral part of the definition.

Sumakuru bigal: holotype, W.P. Maddison, 2016. ZooKeys 614: 87–96. f. 1-11. Ecuador. By courtesy.
Lyssomaneae : - characters used by Simon 1901a.