RETURN TO DATABASE - ALTERNATIVE CLASSIFICATION
SALTICIDAE OF THE WORLD - 0-INDEX of Genera - 01-FOREWORD & KEY to groups of genera! - 02-Comparison - HISTORICAL classification of Aelurilleae - 03-AEURILLINES - 04-AMYCINES- 05-AMYCOIDA VARIA - 06-ASTIAINES - BALLINES - 07-BELIPPINES - 08-CHRYSILLINES - 09-COCALODINES - 10-COLONINES[=Thiodinines] - 11-DENDRYPHANTINES - 12-DIOLENINES - EUODENINES - 13-EUOPHRYINES-PART 0: Introduction-Diversity - 13-EUOPHRYINES-PART 1: Admestina-Donoessus - 14-EUOPHRYINES-PART 2: Echeclus-Pystira - 15-EUOPHRYINES-PART 3: Rhyphelia-Zenodorus - 16-EUPOAINES - 17-EVARCHINES - 18-FOSSILS - 19-HABRONATTINES - 20-HARMOCHIRINES - 21-HELIOPHANINES - 22-HISPONINES - 23-HYLLINES - 24-ICIINES - 25-LAPSIINES - 26-LIGONIPEINES - 27-LYSSOMANINES - 28-MENEMERINES - 29-MYRMARACHNINES - 30-NOTICIINES - 31-PELLENINES - 32-PSEUDICIINES - 33-SIMAETHINES - 34-SITTICINES - 35-SPARTAEINES - 35a-SYNAGELINES - 36-THIRATOSCIRTINAE - 37-YAGINUMAELLINES - 38-YLLENINES - 39-UNCLASSIFIED TEMPORARY
APPENDICES -AP1-Simon's classification - AP2-Petrunkevich synthesis-1928 - AP3-Bonnet's list of subfamilies - AP4-Prˇszynski's revision of subfamilies 1976 - AP5-Prˇszynski - summary of results - AP6-Aelurillus black & gray - AP7-Maddison's views on Salticidae phylogeny 2014 - MS1-Omoedus synonymy

MADDISON'S ALTERNATIVE: Tittle_pg Index Introduction Agoriini Aelurillina Asemoneinae Amycini Amycoida-incertae sedis Astiini Ballini Baviini Bredini Chrysillini Cocalodini Dendryphantina Euophryini-1 Euophryini-2 Euophryini-3 Eupoinae Freyina Gophoini Harmochirina Hasariini Hisponinae Holcolaetina Huriini Itatina Lapsiini Leptorchestini Lyssomaninae Marpissina Mopsini Myrmarachnini Nannenini Neonini Onomastinae Plexippina Salticini Sarindini Scopocirini Simaethina Simonellini Sitticini Spartaeina Synagelina Thiodinini Thiratoscirtina Tisanibini Viciriini Salticidae-incertae sedis

Monograph of the Salticidae (Araneae) of the World 1995-2015. Part I: Introduction to alternative classification of Salticidae.

Jerzy Prˇszyński
09 - COCALODINES GROUP OF GENERA
Version July 1st, 2016.
Sources and permissions of illustrations are displayed in the IInd part of the monograph http://www.peckhamia.com/salticidae/

This chapter contains following genera - see page indicated: type genus Cocalodes
Allococalodes , Cocalodes , Cucudeta , Grayenulla , Sonoita , Sparbambus , Tabuina , Yamangalea .(24 species)

Introduction

COCALODINES contains a few, poorly known, New Guinean genera. It is said that males are characterized by presence of median apophysis on bulbus. Females have small, globular spermathecae, usually located anteriorly, and very thin copulatory ducts. REMARKS: diagnostic drawings of palps in LAPSINES, another poorly know but South American group of genera, show presence of bulbus apophysis. The group is too little known to interpret significance of that structure.

Guide for identification of genera

Gen. Allococalodes Wanless, 1982 (3 species)

Type species Allococalodes alticeps . Drawings below are integral part of the definition.

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Allococalodes alticeps + A. madidus : Wanless 1982a. Bull. brit. Mus. nat. Hist. (Zool.).42 (4): 291-292, ff. 16A-F
+ Maddison, 2009.2021: 5-7, f 10-19. ©Magnolia Press. By courtesy.
Allococalodes cornutus : Wanless 1982a. Bull. brit. Mus. nat. Hist. (Zool.).42 (4): 291-292, ff. 17A-E. By courtesy.

Gen. Cocalodes Pocock, 1897 (12 species) ,

Type species Cocalodes leptopus . Drawings below are integral part of the definition.

Cocalodes sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f.19. By courtesy.

Cocalodes leptopus + C. macellus : Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 275-277, ff. 7A-I, 20A-B
Maddison, 2009: 5, f. 2-9.Magnolia Press. By courtesy .
Cocalodes cygnatus : Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 273-275, ff. 6A-G; 19E-F. By courtesy .
Cocalodes expers : Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 271-273, ff. 5A-C. By courtesy .
Cocalodes innotabilis : Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 288-289, ff. 15A-F, 18A-B. By courtesy .
+
Cocalodes longicornis: Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 277-279, ff. 8A-H; 21A + Australian Museum,
KS48875, photo: M. Zabka. + Maddison, 2009: 5, f. 6-9. .Magnolia Press By courtesy .
+
Cocalodes longipes + C. protervus : Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 284-286, f. 13A-G
+ 279-280, ff. 9A-B. . By courtesy .

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Cocalodes macellus : Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 275-277, ff. 7A-I,20A-B
+ Maddison, 2009: 5, f. 2-9.Magnolia Press. By courtesy .
+
Cocalodes papuanus : Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 267, ff: 1A-B, 2A-H, 3A-E, 18C, E, 19A-B, 21B + Cocalodes papuanus W. P. Maddison et al. (2016): ZooKeys 549: 1–7, f. 9. By courtesy..
Cocalodes thoracicus: Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 280-283, ff. 10A-E, 11A-E, 18D, 20C-D..
Cocalodes turgidus: Wanless 1982. Bull. brit. Mus. nat. Hist. (Zool.)., 42 (4): 283-284, ff. 12A-D, 20E-F. By courtesy .

Gen. Cucudeta Maddison, 2009 ( 3 species) ,

Type species Cucudeta zabkai. Drawings below are integral part of the definition.

Cucudeta zabkai : Maddison 2009: Zootaxa 2021: 15-17, f. 62-63 . (Type species). © Magnolia Press. By courtesy.
Cucudeta gahavisuka : Maddison 2009: Zootaxa 2021: 17-20, f. 83-86. © Magnolia Press. By courtesy.
Cucudeta uzet : Maddison 2009: Zootaxa 2021: 17, f. 79-82. © Magnolia Press. By courtesy.
Cucudeta[?] sp [AUS-1009] : Whyte, O. Crystal Cascades, Nr Cairns. ©Photo I.R. Macaulay + R. Whyte. By courtesy.
Cucudeta[?] sp [AUS-1350] : Whyte, O. Crystal Cascades, Nr Cairns. ©Photo I.R. Macaulay + R. Whyte. By courtesy.

Gen. Cucudeta-cf Whyte, 0 (unpublished)
a long shot - proposal of classification based on presence of plate-like apophysis on bulbus
and embolus atop membranous distal haematodocha

Type species Cucudeta-cf bugeye.

Cucudeta-cf bugeye : Whyte, 0. Australia, P1170109. ©Photo I.R. Macaulay + R. Whyte. By courtesy.
Cucudeta-cf sp [Aus-331] : Whyte, 0. Australia, P1140524iain. ©Photo I.R. Macaulay + R. Whyte. By courtesy.
NOTE: white setae on clypeus, compare with the previous species!
Cucudeta-cf sp [Aus-1928] : Whyte, 0. Australia, P1140524iain. ©Photo I.R. Macaulay + R. Whyte. By courtesy.
NOTE: interpretation of epigyne - only hypothetic!

Gen. Grayenulla Zabka 1992 (8 species)

Type species Grayenulla dejongi.
DIAGNOSIS. Placement of Grayenulla in this group of genera is tentative, based of presence of strong tegular (?) apophysis, usually not very prominent, but fortunately well visible on one of photos of Grayenulla kiwirrkurra (below), resembling somewhat Tabuina and Yamangalea. There are also two tibial apophyses, however, not particularly prominent. Differs from other genera by squat body and short, robust legs, also chelicerae are short, dentition ids fissidentate. Diagnostic drawings below are integral part of definition.

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Grayenulla dejongi + australensis Type species Zabka 1992a. Rec. Austr. Mus., 44: 177-178, figs 10a-e, 11a-d +180-181, figs 15a-e . By courtesy.
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Grayenulla nova + G. spinimana + G. waldockae Zabka 1992a. Rec. Austr. Mus., 44: 181-182, figs 16a-d + 179-180, figs 14a-f. By courtesy.
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Grayenulla wilganea+ G. wishartorum Zabka 1992a. Rec. Austr. Mus., 44: 271-172, figs 2A-F, 5 + 180-181, figs 15a-e .
Grayenulla kiwirrkurra Whyte (in press) Australia: WA, Kiwirrkurra September 2015 Australia's Bush Blitz species discovery program. Internet. ©Photo R. Whyte - by special permission.

Gen. Sonoita Peckham, Peckham, 1903 (1 species)

Type species Sonoita lightfooti. Drawings below are integral part of the definition.

+>
Sonoita lightfooti : Wanless 1985. Bull. brit. Mus. nat. Hist. (Zool.),, 48 (4): 267, f. 11A-L, 13A-B, 14A-B, 16F
+ Wesolowska, Tomasiewicz, 2008. J. Afrotr. Zool., 4: 48-49, f 189-194, Wesolowska, Cumming 2008.
Ann. zool. 58: 212, f 157-163. By courtesy.

Gen. Sparbambus Zhang J., Woon J., Li D., 2006 (1 species)

Type species Sparbambus gombakensis. Drawings below are integral part of the definition.

Sparbambus gombakensis Zhang, Woon, Li 2006. Raff. Bull. Zool., 54(2): 241-244, f 1-11.. By courtesy.

Gen. Tabuina Maddison, 2009 (3 species) ,

Type species Tabuina varirata. Drawings below are integral part of the definition.

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Tabuina varirata + baiteta : Maddison, 2009. Zootaxa 2021: 2021: 11-13, f 44-50 Type Species © Magnolia Press.
Tabuina rufa : Maddison, 2009. Zootaxa: 2021: 13-14, 51-56 © Magnolia Press.By courtesy.

Gen. Yamangalea Maddison, 2009 (2 species) ,

Type species Yamangalea frewana. Drawings below are integral part of the definition.

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Yamangalea frewana + lubinae+ sp [AUS-259] : Type Species Maddison, 2009. Zootaxa 2021:
8-9, f 20-30. © Magnolia Press. + Zabka, 2012. Queensland. © Zootaxa 3176: 62-63, f 19-27. Queensland. ©Photo R. Whyte.
Diagnostic characters of Cocalodeae - according to Simon.